– Selected Publications by Mark J. Wetzel –
[ page update: 2019-I-03 ]


Oligochaete Nomenclator website:

Reynolds, J.W., and M.J. Wetzel. 2018. Nomenclatura Oligochaetologica Editio Secunda – A catalogue of names, descriptions, and type specimens of the Oligochaeta (launched 6 January 2014).

World Checklist of Freshwater Oligochaetes, on-line:

Martin, P., Martinez-Ansemil E., Pinder, A., Timm T. & Wetzel, M.J., 2016 (December 15). World checklist of freshwater Oligochaeta species. World Wide Web electronic publication. Available online at http://fada.biodiversity.be/group/show/12 [date accessed] [this is an on-line update of our 2008 publication {Hydrobiologia, 595, 117-127; full citation listed below}, with support from FADA, the Freshwater Animal Diversity Assessment Initiative]. 

Journal Articles and Peer-Reviewed Publications

Robinson, J.L., M.J. Wetzel, and J.S. Tiemann. 2017. Some phoretic associations of macroinvertebrates on transplanted federally endangered freshwater mussels. Northeastern Naturalist 24(4): N29-N34.

Schenková, J., M.J. Wetzel, E. Martínez-Ansemil, and P. Martin. 2016.  Editorial. 13th International Symposium on Aquatic Oligochaeta, Brno, Czech Republic, 7–11 September, 2015. Biologia 71(1): 1-2.  DOI: 10.1515/biolog-2016-0016.

Rota, E., S. Martinsson, M. Bartoli, A. Beylich, U. Graefe, A. Laini, M.J. Wetzel, and C. Erséus. 2016. Mitochondrial evidence supports a Nearctic origin for the spreading limicolous earthworm Sparganophilus tamesis Benham, 1892 (Clitellata, Sparganophilidae). Contributions to Zoology 85(1): 113-119.

Schenková, J., M.J. Wetzel, E. Martinéz-Ansemil, and P. Martin. 2016. Editorial. 13th International Symposium on Aquatic Oligochaeta, Brno Czech Republic, 7–11 September, 2015. Biologia 71(1): 1-2. 

Martinsson, S., Y. Cui, P.J. Martin, A. Pinder, K. Quinlan, M.J. Wetzel & C. Erséus. 2015. DNA-barcoding of invasive European earthworms (Clitellata: Lumbricidae) in south-western Australia. Biological Invasions 17(9): 2527-2532.

Pinder, A., N. Arslan, and M. Wetzel (eds.). 2014. Proceedings of the 12th International Symposium on Aquatic Oligochaeta, convened in Fremantle, Western Australia, 10-13 September 2012. Zoosymposia, Vol. 9(1). 102 pp.  ISBN 978-1-77557-410-1 (on-line version, open access; paper version also available). 12 June.

Pinder, A., N. Arslan, and M. Wetzel (eds.). 2014. Preface. Proceedings of the 12th International Symposium on Aquatic Oligochaeta. Zoosymposia 9(1): 4-5 (full citation, above).

Reynolds, J.W., and M.J. Wetzel. 2014.  A checklist by counties of earthworms (Oligochaeta: Acanthodrilidae, Lumbridicae, Megascolecidae, and Sparganophilidae) in Michigan, USA.  Megadrilogica 17(5): 51-72. 

Major, K., D.J. Soucek, R. Giordano, M.J. Wetzel, and F. Soto-Adames. 2013.  The common ecotoxicology laboratory strain of Hyalella azteca is genetically distinct from most wild strains sampled in eastern North America.  Environmental Toxicology and Chemistry, Vol. 32(11): 2637–2647.

Arslan, N., and M. J. Wetzel. 2012. Preface.  Pp. iii-vi, In: Colack, Ercument; Arslan, Naime; Wetzel, Mark J.; Timm, Tarmo; Wang, Hong-Zu; Pinder, Adrian; and Schenkova, Jana, editors. Aquatic Oligochaete Biology XI. Proceedings of the 11th International Symposium on Aquatic Oligochaeta held in Alanya, Antalya, Turkey, 5010 October 2009. viii + 145 pp.  Turkish Journal of Zoology 36(1).  [but note that the preface...was not included on this website]

Reynolds, J.W., and M.J. Wetzel. 2012. Terrestrial Oligochaeta (Annelida: Clitellata) in North America, including Mexico, Puerto Rico, Hawaii, and Bermuda. III. Megadrilogica 15(8): 191-209.

Stribling, J., B. Sweeney, J. Morse, G. Corkum, G. Lester, S. Miller, R. Mitchell, B. Poulton, S. Strachan, and M. Wetzel. 2012. Taxonomic certification versus the scientific method”: a rebuttal of Rogers (2012).  Zootaxa 3359: 65-68.  {and be sure to visit the erratum for the above paper, published in a subsequent issue [Zootaxa 3383: 14.]}

Krysko, K.L., M.C. Granatosky, J.C. Anderson, M.J. Wetzel, and D.J. Klemm. 2012. Nerodia fasciata (Southern Watersnake): Ectoparasites. Herpetological Review 43: 347-348.

Reynolds, J.W., and M.J. Wetzel. 2011. The earthworms (Oligochaeta: Acanthodrilidae, Glossoscolecidae, Komarekionidae, Lumbridicae, Megascolecidae, and Sparganophilidae) of Illinois, USA.  Megadrilogica 15(4): 35-67. 

Reynolds, J.W., and M.J. Wetzel. 2008. Terrestrial Oligochaeta (Annelida: Clitellata) in North America, including Mexico, Puerto Rico, Hawaii, and Bermuda. Megadrilogica 12 (12): 157-208.

Wang, H.-Z., M.J. Wetzel, A.M. Pinder, P.F.M. Verdonschot, and N. Arslan (eds.). 2008. Aquatic Oligochaete Biology X. Proceedings of the 10th International Symposium on Aquatic Oligochaeta, held at the Institute of Hydrobiology, Chinese Academy of Sciences in Wuhan, China, 16-21 October, 2006. Acta Hydrobiologica Sinica 31 (supplement). viii + 171 pp. ISSN 1000-3207.

Wang, H.-Z., and M.J. Wetzel, 2008. Preface. Pp. i-ii, In: Wang, H.-Z. et al. (eds.). Aquatic Oligochaete Biology X. Proceedings of the 10th International Symposium on Aquatic Oligochaeta. Acta Hydrobiologica Sinica 31 (suppl.).

Erséus, C., M.J. Wetzel, and L. Gustavsson. 2008. ICZN rules--a farewell to Tubificidae (Annelida, Clitellata). Zootaxa 1744: 66-68.

Readel, A.M., C.A. Phillips, and M.J. Wetzel. 2008. Leech parasitism in a turtle assemblage: effects of host and environmental characteristics. Copeia 2008(1): 227-233.

Martin, P., E. Martínez-Ansemil, A. Pinder, T. Timm, and M.J. Wetzel. 2008. Global diversity of oligochaetous clitellates ("Oligochaeta"; Clitellata) in freshwater. Pp. 117-127, In E. V. Balian, C. Lévêque, H. Segers & K. Martens, eds. Freshwater Animal Diversity Assessment. Hydrobiologia 595(1). Access this paper as an on-line (html) file HERE.

Favret, C., K.S. Cummings, R.J. McGinley, E.J. Heske, K.P. Johnson, C.A. Phillips, L.R. Phillippe, M.E. Retzer, C.A. Taylor, & M.J. Wetzel. 2007. Profiling natural history collections: a method for quantitative and comparative health assessment. Collection Forum 22(1-2): 53-65.

Wetzel, M.J., and M.A.P. Morgan. 2007. Aquatic microdrile oligochaetes (Annelida, Clitellata) of Great Smoky Mountains National Park, North Carolina and Tennessee. Pp. 153-158, In: P. Cox, (Ed.). The Great Smoky Mountains National Park All Taxa Biodiversity Inventory: A Search for Species in Our Own Backyard. Southeastern Naturalist 6 (Special Issue 1). 238 pp.

Nijboer, R.C., M.J. Wetzel, and P.F.M. Verdonschot. 2004. Diversity and distribution of Tubificidae, Naididae, and Lumbriculidae (Annelida: Oligochaeta) in the Netherlands: an evaluation of twenty years of monitoring data. Hydrobiologia 520: 127-141.

Reynolds, J.W., and M.J. Wetzel. 2004. The terrestrial Oligochaeta (Annelida: Clitellata) of North America north of Mexico. Megadrilogica 9(11): 71-98.

Swayne, H., M. Day, and M.J. Wetzel. 2004. Limnodrilus hoffmeisteri (Annelida: Oligochaeta: Tubificidae) in Pop's Cave, Wisconsin, USA. Journal of Cave and Karst Studies 66(1): 28-31.

Kathman, R.D., and M.J. Wetzel. 2003. Allonais inaequalis (Annelida: Oligochaeta: Tubificidae) in North America. Proceedings of the Biological Society of Washington 116(3): 548-556.

Wetzel, M.J., and P. Morgan. 2002. Aquatic Oligochaeta (Annelida, Clitellata) of the Great Smoky Mountains National Park, North Carolina and Tennessee. Southeastern Biology 49(4): 389-392.

Wetzel, M.J., and S.J. Taylor. 2001. First records of freshwater oligochaetes (Annelida, Clitellata) from caves in Illinois and Missouri, USA. Journal of Cave and Karst Studies 63(3): 99-104. Available in PDF: < http://www.caves.org/pub/journal/PDF/V63/v63n3-Wetzel.pdf >

Britton, D.K., and M.J. Wetzel. 1999. First records for freshwater oligochaetes (Annelida, Oligochaeta, Naididae) in the Hawaiian Islands with notes on their association with an alien snail (Mollusca, Gastropoda, Ampullariidae). Occasional Papers of the Bishop Museum - Records of the Hawaii Biological Survey for 1998 59(Part 2): 39-42.

Wetzel, M.J., G. Oberlin, and D.W. Blinn. 1999. The aquatic Oligochaeta (Annelida: Clitellata) of Montezuma Well, Arizona: a near thermally constant limnocrene. Southwestern Naturalist 44(4): 514-518.

Webb, D.W., M.J. Wetzel, P. C. Reed, L. R. Phillippe, and T. C. Young. 1998. The macroinvertebrate biodiversity, water quality, and hydrogeology of ten karst springs in the Salem Plateau Section of Illinois, USA. Pp. 39-48, In: L. Botosaneanu (ed.). Studies in Crenobiology - The biology of springs and springbrooks. Backhuys Publishers, Leiden, The Netherlands. 261 pp. ISBN 90-73348-04-8.

Webb, D.W., M.J. Wetzel, P.C. Reed, L.R. Phillippe, and M.A. Harris. 1995. Aquatic biodiversity in Illinois springs. Journal of the Kansas Entomological Society 68(2)suppl.: 93-107.

Wetzel, M.J. 1992. Aquatic Annelida of Illinois: Introduction and checklist of species. Transactions of the Illinois State Academy of Science 85(1 and 2): 87-101.

Wetzel, M.J. 1987. Limnodrilus tortilipenis, a new North American species of freshwater Tubificidae (Annelida: Clitellata: Oligochaeta). Proceedings of the Biological Society of Washington 100(1): 182-185.

Brinkhurst, R.O., and M.J. Wetzel. 1984. Aquatic Oligochaeta of the world: Supplement. A catalogue of new freshwater species, descriptions, and revisions. Canadian Technical Report of Hydrography and Ocean Sciences No. 44. v + 101 pp.

Wetzel, M.J. 1982a. Kansas leeches (Annelida: Hirudinea) with notes on distribution and ecology. II. Technical Publications of the State Biological Survey of Kansas 12: 105-111.

Wetzel, M.J. 1982b. Aquatic Oligochaeta (Annelida: Clitellata) in Kansas, with notes on their distribution and ecology. Technical Publications of the State Biological Survey of Kansas 12: 112-130.

Seagle, H.H., Jr., and M.J. Wetzel. 1982. Range extension of Barbidrilus paucisetus Loden and Locy (Oligochaeta: Enchytraeidae). Freshwater Invertebrate Biology 1(4): 52-53.

Wetzel, M.J. 1980. Limnodrilus rubripenis Loden and Psammoryctides californianus Brinkhurst, two aquatic worms (Annelida: Oligochaeta: Tubificidae) new to Illinois. Transactions of the Illinois State Academy of Science 73(3): 36-38.

Klemm, D.J., D.G. Huggins, and M.J. Wetzel. 1979. Kansas leeches (Annelida: Hirudinea) with notes on distribution and ecology. Technical Publications of the State Biological Survey of Kansas 8: 38-46.


Memoriams

Timm, T., and M.J. Wetzel. 2014. In memoriam: Tamara L. Poddubnaja (1930–2011) – Oligochaetologist. Zoosymposia 9(1): 6-9. In: Pinder, A., N. Arslan, and M. Wetzel (eds.). 2014. Proceedings of the 12th International Symposium on Aquatic Oligochaeta, convened in Fremantle, Western Australia, 10-13 September 2012. Zoosymposia, Vol. 9(1). 102 pp.  ISBN 978-1-77557-410-1 (on-line version, open access; paper version also available). 12 June.

Wetzel, M.J. 2008. In memoriam: Charlie D. Drewes (1946-2005). Pp. 10-17, In: Wang, H.-Z. et al. (eds.). Aquatic Oligochaete Biology X. Proceedings of the 10th International Symposium on Aquatic Oligochaeta. Acta Hydrobiologica Sinica 31 (suppl.).

Wetzel, M.J. 2008. In memoriam: Michael Robert Milligan (1951-2005). Pp. 18-20, In: Wang, H.-Z. et al. (eds.). Aquatic Oligochaete Biology X. Proceedings of the 10th International Symposium on Aquatic Oligochaeta. Acta Hydrobiologica Sinica 31 (suppl.).

Wetzel, M.J. 2006. In memoriam: Professor Walter James Harman, PhD (1928-2002). Hydrobiologia 564(1): 1-4. [also, Pp. 1-4, In: Verdonschot, P.F.M., H. Wang, A. Pinder, and R. Nijboer (eds). 2006. Aquatic Oligochaete Biology IX. Selected papers from the 9th Symposium on Aquatic Oligochaeta, 6-10 October 2003, Wageningen, The Netherlands. Developments in Hydrobiology 186. Published by Springer, Dordrecht, The Netherlands. ix + 222 pp. ISBN 1-4020-4781-9.


Other Publications

Reynolds, J.W., and M.J. Wetzel. 2018. Nomenclatura Oligochaetologica Editio Secunda – A catalogue of names, descriptions, and type specimens of the Oligochaeta (launched 6 January 2014).

Wetzel, M.J., S.V. Fend, K.A. Coates, R.D. Kathman, and S.R. Gelder. 2009. Taxonomy, systematics, and ecology of the freshwater oligochaetes and branchiobdellidans (Annelida, Clitellata) of North America. A workbook. 1 November. vi + 280 pp. + 4 color plates. Copyright (c) 2009, November. {The overall scope of this workbook is the North American continent; this serves as a compliment to the most recent guide published by R.D. Kathman and R. Brinkhurst in 1998; although this workbook is now out of print / no longer available, an updated and expanded version is being prepared.}

Wetzel, M.J., and P. Morgan. 2002. Worms: Segments, Hairs, Maybe Gills -- Oh My! ATBI Quarterly 3(1): 6-7 [Winter 2002 issue]. Available in PDF, at:  < http://www.discoverlife.org/pa/pu/ATBI_Quarterly/atbi_quarterly_winter_2002.pdf >

Wetzel, M.J., and D.J. Watermolen. 1995. Horsehair worms in Illinois. Illinois Natural History Survey Reports No. 332: 4; 10.

Wetzel, M.J. Deformities in aquatic macroinvertebrates. 1994. Illinois Natural History Survey Reports No. 327: 6-7.

Webb, D.W., M.J. Wetzel, L.R. Phillippe, and P.C. Reed. 1993. Springs of Illinois. Illinois Natural History Survey Reports No. 323: 2-3.


Papers in preparation or in press:

Several papers (with collaborators) on distribution records for 1) earthworms of Illinois (revisited); 2) records of freshwater oligochaetes new to North America; 3) freshwater oligochaetes and earthworms in the Huron Mountain area, U.P. Michigan (2 papers); 4) earthworms in North America (fourth update).  


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